A proposal for a project (3rd year, master)

Adding Time to Discrete Models of Genetic Regulatory Networks

In the book of Thomas and D'Ari, Biological Feedback, the author introduce an asynchronous discrete models for describing the dynamics of genetic regulatory network. In these models the dynamics is based on the following principles:

  • The state space is discretized (gene is "on" or "off", and a protein product is present or absent (the discretization can be refined to include various ranges of product levels;

  • The interaction between genes and products are given qualitatively by diagrams with arrows which mean something like gene X encourages the production of product x, or more complex dependencies between several variables;


  • The dynamics is based on delays: when a gene X is "on" than after some time product x will become present


  • These models are asynchronous and qualitative: they do not suppose any relations between the delays of different reactions and hence the "global" automaton representing the system may have many behaviors, each corresponding to a different choice of the delay values.

    The project intends to extend the model with timing information on the reactions and use timed automata to model the whole network. Timing constraints will reduce the set of behaviors by eliminating behaviors the violate these constraints. The modeling will be inspired by the work of Maler and Pnueli (95) on quantitative modeling of delays in digital circuits.

    The work will include understanding of the kinetic logic model of Thomas and D'Ari, learning the principles of timed automata and circuit delays and then defining new models and studying their dynamical properties, possibly using existing verification tools. Some applications in synthetic biology will be considered.

    References:

  • The book Biological Feedback which is available online

  • O. Maler and A. Pnueli paper on asynchronous circuits and timed automata

  • A recent attempt to add time to the models by Siebert and Bockmayr